(after Horak and Brown, 1991)
Introduction
The Tortricinae constitute a polyphyletic group, but there are no unequivocal synapomorphies to either associate particular tribes more closely within the subfamily or to refer them elsewhere.
DIAGNOSIS
Diagnostically important characters are given in italics
scale arrangement male antenna
wing venation (showing cubital pecten) (only present in Sparganothini)
male genitalia Tortricinae (examples)
male genitalia (explaining terms)
aedeagus
female genitalia (explaining terms)
Antenna nearly always with two rings of scales per flagellar segment, often ciliate. Forewing often with costal fold. Hindwing only exceptionally with cubital pecten. Male genitalia variable, but juxta and aedeagus nearly always articulated, not strongly fused; aedeagus with coecum usually developed. Female genitalia: sterigma connected with ventral arms of anterior apophyses (connection reduced in Cnephasiini); usually one signum, only exceptionally two. Majority of pheromones based on a 14-carbon chain.