(after Horak and Brown, 1991)
Introduction
Cochylini (examples of male moths)
Cochylini (about 700 species) are represented in all faunal regions, with the highest diversity in the drier southern parts of the Palaearctic and only a single species known from Australia. The mostly oligophagous Cochylini usually feed internally in flower heads, seed capsules, stalks and roots, rarely in rolled leaves of mainly herbaceous plants in a wide range of families.
DIAGNOSIS
Diagnostically important characters are given in italics
Adult
venation Eupoecilia angustana (explaining terms)
male genitalia Cochylini (explaining terms)
female genitalia Cochylini (explaining terms)
Labial palpus porrect, second segment expanded apically. Forewing with termen oblique and never deeply sinuate ; often with refractive scales; pattern usually based on two inwardly oblique transverse fasciae, with a conspicuous oblique mark at mid-dorsum; CuP absent or strongly reduced, CuA2 from 2/3 cell or more distally. Male genitalia: uncus often atrophied; gnathos reduced or atrophied; aedeagus usually very large, often with numerous cornuti. Female genitalia: bursa rarely separated into ductus and corpus ; usually with accessory bursa; with often heavily sclerotized antrum; bursa with numerous spines and/or various sclerites, never with well defined signum.
Larva
setal map (showing position of setae. Note: example is not a cochyline larva)
Anal fork often present, crochets uniordinal except in some Trachysmia Guenée; D1 and SD1 on same pinaculum on A9, but D2 setae sometimes on separate pinacula; SD2 at least sometimes on separated pinacula on A1-A7; L group usually bisetose on A9, but trisetose in some Trachysmia ; SV group on A1, 2, 7, 8 and 9 is very variable, from 3:3:3:2:2 to 2:2:2:1:1; V1 on A9 further apart than on A8.
Pupa
Stenodes spp., Aethes spp.: frons sometimes with large ventral thorn, alar furrow a shallow depression; A2-A7 with two, A8 and A9 with one row of spines (Stenodes alternana (Stephens) with cephalic rows as bands of minute spines), and A10 always with several spines, often nearly encircling anal rise; no cremaster, but two spines often enlarged into two subdorsal thorns, and smaller thorns often present on each side of anal rise; A10 with weak hooked setae on apex and close to lateral thorns, but not with distinct setae on anal rise.
Genera including pests (in Europe and Asia)
Eupoecilia
Cochylis